Key to Wisconsin Freshwater Snails

Developed by Kathryn E. Perez and Greg Sandland, based on previous key by J.B. Burch 1980.

Copyright notes: Unless otherwise noted, all images are copyright UWL and taken by Chris Lynum. Shell drawings by Brittany Harried, modeled on images in Burch 1980.

Use of key: Taxonomy follows Turgeon et al. 1998, unless revisionary taxonomic work has been published since 1998. References provided at the end of this page. Also note that key is for adult individuals of extant species.  Fossil species can be accessed via other keys. Snail vocabulary terms can be found in the glossary. Use of the key: The key begins with a few couplets to bring you to family of snails. Then separate couplets are for each family. If you already know family, go directly to that point. All shell photographs with a ruler have lines with an interval of 1 mm.

Additional useful pages: Key to the egg masses of WI snails, Comparison of difficult snails, Size comparison of juvenile Chinese mystery snails to other native snails.

Key to the freshwater snail families of Wisconsin

1a.  Animal with an operculum (NOTE - opercula may be missing in some specimens due to preservation methods, age of specimen, etc; therefore it is important to use other morphological characters in suspected cases) (2)

1b.  Animal without an operculum (7)

2a.  Shell small (diameter up to 5 mm), spire generally depressed, some species with carina, operculum multispiralValvatidae

2b.  Shell small to large, spire depressed to elongate, operculum multispiral, paucispiral, or concentric (3)

2a. 

3a.  Operculum multispiral or paucispiral, distal margins not concentric (4)

3b.  Operculum concentric (6)

3a.   File:Marstonia comalensis operculum.png 3b.File:Filopaludina martensi operculum.png

4a. Adult shell < 5 mm in length (5)

4b.  Adult shell medium, to large (> 20 mm in length)...Pleuroceridae

5a.  Shell high spired, turriform; head-foot subdivided on each side by a longitudinal groove…Pomatiopsidae

5b.  Shell high spired to depressed; head-foot region lacking a longitudinal groove…Hydrobiidae

5a.      5b.

6a.   Shells of adults up to 15 mm in length; Operculum calcareous and concentricBithyniidae, Bithynia tentaculata

6b.  Shells of adults medium to large (> 20 mm); operculum corneousViviparidae

6a.         6b.

7a.  Shell coiled (8)

7b.  Shell not coiled; cone-shaped (freshwater limpets)…Ancylidae

7a.      7b.

8a.  Animal shell is dextralLymnaeidae

8b.  Animal shell sinistral (9)

8a.      8b.

9a.  Shell with raised spire…Physidae

9b. Shell discoidal with sunken spire…Planorbidae

9a.     9b.

 

 

Planorbidae

The unique flattened shape of most planorbids requires some attention to terminology before keying. The shells of planorbids are sinistral, with the aperture opening to the left. When shells are held with the aperture toward the viewer and to the left, the spire is on top and the umbilicus is on the bottom.

1a. Shell small, that of adults < 8 mm in diameter…… Gyraulus and Promenetus

1b. Shell larger, that of adults > 8 mm and up to or more than 30 mm in diameter... (7).

2a. Spire pit relatively wide.. Gyraulus (3)

2b. Spire pit deep and narrow… Promenetus (6)

2a.         2b.

 

3a. Shell costate, having large surface sculpture... Gyraulus crista  

3b. Shell not costate (4)

3a. Gyraulus crista        3b. Gyraulus deflectus

4a. Adult shells 4-7 mm in diameter, variable, with the body whorl not evenly rounded or with a peripheral keel or with a hirsute periostracum or a malleated surface, or these features combined... Gyraulus deflectus

4b. Adult shells 3-5 mm in diameter, variable with the body whorl evenly rounded or with upper lateral surface slightly flattered; without a peripheral keel, hirsute periostracum or malleated surface (5)

4a.         4b.

5a. Shell relatively high... Gyraulus hornensis

5b. Shell relatively flattened, whitish or yellowish, semi-transparent, entirely or nearly planispiral, appearing almost the same from both sides... Gyraulus circumstriatus

5a.            5b.

6a. Shell with carinate periphery, relative height of body whorl nearly equal from one side to the other... Promenetus exacuous

6b. Shell with rounded periphery, relative height of body whorl increasing toward the aperture... Promenetus umbilicatellus

6a.        6b.

7a Inside aperture or body whorl with “teeth” or lamellae... Planorbella armigera.

7b. Aperture or body whorl without teeth or lamellae (8)

7a.         7b.

8a. Shell spire strongly inverted, with a more or less conical depression; spire side of whorl with or without a strong keel....Helisoma anceps

8b. Shell spire not strongly inverted, with a shallow depression, no depression or exverted (raised above body whorl); spire side of body whorl rounded or angular... Planorbella (9)

8a.      8b.

9a. Body whorl at shell aperture campanulate (flared)... Planorbella campanulata

9b. Body whorl at shell aperture straight, not campanulate. Shell surface usually dull, rough in texture, with raised transverse thread-like striae. (10)

9a.         9b.

10a. Carinae or strong angulations present on the outer edges of both the right (umbilical) and left (spire) side of the body whorl of the shell. Shells small, those of adults less than 18 mm in greatest diameter, spire flat, not inverted or sunken into a bowl-like depression. (11)

10b. Carinae absent, although a rather strong angulation might be present on the upper surface of the body whorl of the spire. (12)

10a.        10b.

11a. Shells small, those of adults less than 18 mm in greatest diameter, spire flat, not inverted or sunken into a bowl-like depression, lines of growth more moderate, flattened...Planorbella truncata

11b. Shells large, very wide whorls, pronounced longitudinal sculpture, these lines of growth are distant, elevated, and conspicuous, giving surface a rough appearance...Planorbella binneyi

11a.         11b. Planorbella binneyi

 12a. Inverted portion of the shell spire relatively wide, concavely smooth-sided and bowl-like....Planorbella pilsbryi

12b. Inverted portion of shell spire narrower, generally not smooth-sided or bowl like....Planorbella trivolvis

12a.         12b.

Pleuroceridae

1a. Anterior or “basal” end of aperture prolonged into a short canal, producing an auger-shaped base to the shell. up to 10 whorls, sometimes whorls lighter in color on the upper (closer to spire) portion of whorl. Aperture ~1/3 height of shell... Pleurocera acuta

1b. Anterior or “basal” end of aperture not channeled or auger-shaped. ~5-7 whorls, uniform in color. Aperture ~1/5 height of shell... Elimia livescens

1a.             1b.

 

Valvatidae

1a. Shell lacking postnuclear spiral carinae or angulations (2)

1b. Shell with one to three postnuclear spiral carinae or angulations (4)

2a. Shell depressed-turbinate, with spire slightly elevated above the body whorl (3)

2b. Shell high-turbinate or subconical, spire markedly elevated (5)

3a. Shell diameter > 5 mm, axial striae lamellate; luster of shell dull... Valvata lewisi

3b. Shell diameter < 5 mm, color of apical whorls usually dull purple, or violet, or pink. Luster of shell dull... Valvata perdepressa

3a.             3b.

4a. Shell depressed, with spire barely visible above body whorl shell with 2 spiral carina or angulations, shoulder of the body whorl sloping downward from the dorsal carina to the suture... Valvata bicarinata

4b. Shoulder of the body whorl sloping upward from the dorsal carina nearly to suture, then turning downward... Valvata winnebagoensis

4a.             4b.

5a. Umbilicus narrow, no interior whorls visible, apex of shell flattened appears truncated... Valvata piscinalis

5b. Umbilicus wider, may see interior whorls, apex of shell acute, shell diameter >5 mm, shell often pale green in color, may have fine axial striae... Valvata sincera

5a.             5b.

Viviparidae

1a. Shell large, adults >35 mm and up to 55 mm in length, shell relatively thin, whorls not shouldered... Cipangopaludina (2)

1b. Shell medium to large, generally less than 35 mm in length, but if large, the shell is thick and ponderous, and the whorls are generally shouldered (the body whorl is slightly flattened near where it intersects with the previous whorl) (3)

1a.                1b.

2a. Shell with acute spire, protoconch is elevated, and usually with spiral angulations or low carinae on the whorls; not malleated... Cipangopaludina japonica

2b. Shell with obtuse spire, protoconch flat, and without spiral angulations or low carinae; generally with surface malleations, may have several spiral lirae on body whorl, shell tends to be shouldered... Cipangopaludina chinensis

2a.              2b.

3a. Operculum concentric, but with a spiral nucleus; whorls commonly with a spiral subsutural sulcus... Lioplax sulculosa

3b. Operculum entirely concentric, including its nucleus; whorls without spiral sulci (4)

3a.             3b.

4a. Shell with or without spiral color bands, width and length of aperture usually nearly equal, making it round, or nearly so... Viviparus (5)

4b. Shell without spiral color bands; length of aperture noticeably greater than width... Campeloma. (7)

5a. Shell dark yellowish-green to (usually) dark olive-green, without spiral color bands; shell broadly ovate, whorls globosely rounded, spire obtuse... Viviparus intertextus.

5b.Shell pale olive-green to olive-brown; with or without spiral color bands, ovate but not broadly so, whorls flattened to well rounded, but not globosely rounded, spire relatively acute (6)

5a.             5b.

6a. Shell yellowish-brown or olive-brown; color bands, when present, three in number, shell rather heavy, whorls often flat-sided... Viviparus subpurpureus

6b. Shell yellowish-green or olive-green; color bands, when present, usually four in number, shell relatively thin, but sturdy, whorls usually well rounded... Viviparus georgianus

6a.             6b.

7a. Shell large, heavy, and ponderous... Campeloma crassulum

7b. Shell medium or a little larger, relatively thin to strong, but not very large or heavy and ponderous... Campeloma decisum

7a.                     7b.

Ancylidae

General - Low, obtuse cap-shaped shells; often small in size with the shell apex offset to the right; long thin tentacles.  Freshwater limpets

1a.  Shell typically elevated with fine radial striae on apex.  Aperture narrow to broadly ovate (2)

1b.  Shell typically depressed with no trace of radial striae.  Aperture typically ovate to subcircular, smooth with small raised riblets on the anterior slope.  Apex behind the center of the shell...Laevapex fuscus

1a.             1b.

2a.  Shell to 7mm in length.  Shell is robust, elevated and the aperture is elliptical.  Apex in midline or slightly offset to the right.  Anterior slope convex, posterior slope slightly concave.  Calcareous material is often thick inside the shell... Ferrissia rivularis

2b.  Shell depressed or moderately elevated; rarely exceeding 3.5 mm in length.  With or without a shelf-like septum across the posterior part of the aperture.  When non-septate, aperture is oval in appearance and wider anteriorly.  When septum is present, the shell is evenly elliptical…Ferrissia fragilis

2a.             2b.

 

Hydrobiidae

1a.  Columella and parietal wall not thickened (2)

1b.  Columella usually thickened (8)

1a.     1b.

2a.  Shell rimate or imperforated (3)

2b.  Shell umbilicated (4)

2a.         2b.

3a.  Top of spire truncated; first several whorls coiled in the same plane... Probythinella emarginata

3b.  Top of spire not truncated, first several spire whorls coiled in a descending spiral… Hoyia sheldoni

3a.     3b.

4a.  Aperture broadly ovate to broadly ellipsoid… Cincinnatia integra

4b.  Aperture simply ovate (5)

4a.             4b.

5a.  Shell elongately conic (6)

5b.  Shell typically ovately conic (7)

5a.         5b.

6a. Whorls rounded, almost shouldered, sutures more deeply incised. Spire is rounded, aperture is oval, apertural lip is reflected only slightly at base of lip. Fontigens nickliniana

6b. Compared to Fontigens nickliniana, whorls are less rounded, sutures less incised, spire is more pointed, aperture is not perfectly oval, draw up to a point into almost a teardrop shape. Aperture is reflected around most of the lip except at the very top. Often a brownish-reddish rim around the lip. U.S. populations will have brood pouch with developed embryos with eyes. Potamopyrgus antipodarum

                      6a.                         6b.new zealand mud snail

7a.  Protoconch punctate Pyrgulopsis lustrica

7b.  Protoconch smooth (8)

8a.  Nuclear whorl on shell relatively small (0.29-0.36mm  in diameter), slightly protruding…Lyrogyrus*

8b.  Nuclear whorl of shell relatively large (0.38-0.48mm in diameter) and flattened… Amnicola limosa

8a.             8b.

9a.  Adult shell > 6mm in length… Birgella subglobosa

9b.  Adult shell < 6mm in length (10)

9a.             9b.

10a.  Shell with a distinct umbilical chink; columella thickened… Somatogyrus depressus

10b.  Shell imperforate or rimate; columella with heavy callus forming a thickened continuous peristome Somatogyrus tryoni

10a.            10b.

Note: *Although two species of Lyrogyrus (L. pilsbryi and L. walker) were reported from Wisconsin, specific and reliable features used to distinguishing these snails to species are not available.

 

Lymnaeidae 

General - Broad, flattened triangular tentacles.  Dextral shells that are coiled and tend to be elongated.  Adult shell with narrow or globose body whorl, but if globose, the shell is well sculptured with small spiral striations.

1a.  Shell attenuate, very narrow almost needlelike… Acella haldemani

1b.  Shell thicker, not especially narrow (2)

1a.              1b.

2a.  Shell succiniform (thin and fragile) with large oval aperture and body whorl, and small spire; surface sculptured with raised spiral periostracal threads… Pseudosuccinea columella

2b.  Shell not succiniform, aperture may or may not be large and oval, but if so, the shell is not thin or fragile, and is not sculptured with periostracal threads (3)

2a.              2b.

3a.  Shell large, that of adults > 35 mm in length; shell with a wider, expanded elongately oval to globose body whorl (4)

3b.  Shell smaller, that of adults < 35 mm in length; shell with a relatively narrow body whorl (5)

4a.  Shell with a narrow pointed spire with a large, subglobose body whorl... Lymnaea stagnalis

4b.  Shell with a relatively wide, elongated spire; whorls not shouldered… Bulimnaea megasoma  

4a.      4b.

5a.  Adult shell small (generally < 13 mm); typically lacking spiral sculpture.  Columella generally lacking twist (6)

5b.  Adult shell medium to large, generally > 13 mm in length; surface sculpted with small spiral striations; columella usually with a well-developed twist (11)

6a.  Adult shells with approximately 5 whorls, very small (< 7 mm in length) (7)

6b.  Adult shells larger (> 8 mm in length), shell spire broad to narrow, but in shells with narrow spires, the spire length is not much greater than the aperture length (8)

7a.  Whorls well-rounded, aperture roundly ovate Fossaria parva

7b.  Whorls distinctly shouldered, aperture elongately ovate or elliptical… Fossaria dalli

7a.             7b.  

8a. Whorls regularly or irregularly increasing in size, terminating in an elongate-ovate, sometimes narrow body whorl.  Pointed spire, compressed body whorl, and elongated shouldered aperture.   Inner lip is appressed to the body whorl about the middle of the aperture… Fossaria obrussa

8b.  Shell not as above (9)

8a.             8b.

9a.  Spire of shell is long and the whorls are flatly rounded; very deep suture which is almost channeled in some specimens causing the whorls to become turban-shaped…..Fossaria exigua

9b.  Suture not as pronounced (10)

9a.                9b.

10a.  Shell relatively slender; aperture more oval… Fossaria peninsulae

10b.  Shell also slender, but aperture is more elongated; impressed inner lip where it joins the parietal wall Fossaria modicella

10a.             10b.

11a.  Possess an elongated, rather narrow brown shell (12)

11b.  Possess broader, light-colored shells with subglobose body whorls (14)

11a.             11b.

12a.  Shell quite narrow, flattened elongated whorls Stagnicola exilis

12b.  Shell more elongately conic, rounded body whorl (13)  

12a.              12b.

13a.  Spire whorls long and acutely pointed; sutures deeply impressed; aperture roundly ovate Stagnicola elodes

13b.  Spire whorls short and blunt; sutures not particularly indented; aperture elongately ovate… Stagnicola winnebagoensis

13a.             13b.

14a.  Spire as long as aperture, forming a narrowly acute pyramid (15)

14b.  Spire somewhat shorter that aperture, forming a wide short pyramid (16)

15a.  Inner lip triangular, tightly appressed to columella; axis twisted and plaited... Stagnicola catescopium

15b.  Inner lip forming flat, smooth shelf-like projection over columella; without plait... Stagnicola emarginata

15a.             15b.

16a.  Spire acute, 1/3 the length of shell; aperture roundly ovate Stagnicola walkeriana

16b.  Spire depressed, broad 2/5 the length of shell; aperture elongately ovate… Stagnicola woodruffi

16a.             16b.

Note: Two other species Stagnicola (S. apicina and S. caperata ) have been reported from Wisconsin; however specific and reliable features for distinguishing these snails to species are not available.

 

Physidae

General - Members of this group can be distinguished from members of the Lymnaeidae based on the fact that they have sinistral (left-coiled) shells. 

1a.  Shell elongate, surface glossy (2)

1b.  Shell subglobose with a short spire (3)

1a.     1b.

2a.  Shell nearly spindle-shaped, spire long… Aplexa elongata 

2b.  Shell not as elongated, apex rounded… Physa fontinalis (image by Francisco Welter Schultes)

2a.     2b.

3a.  Shell whorls are convex with deep sutures; spire is pointed; can reach 15 mm at maturity… Physella acuta

3b.  Shell whorls have more flattened sutures; spire is conical; can reach 24 mm at maturity… Physella gyrina

3a.             3b.

Note: Physid shell features can be extremely variable among habitats and this has led to much taxonomic confusion in the past.  The key presented here is based on recent physid phylogenies and descriptions by Pip and Franck (2008), Wethington and Lydeard (2007), Dillon and Wethington (2006), and Burch (1980).

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Acknowledgments

This project was funded by the Wisconsin DNR to assist with documentation of aquatic invasive snails and the native fauna of Wisconsin. Thanks go to UWL students Kyle Fischer and Brittany Harried for key testing. Brittany Harried contributed drawings of snails. The University of Michigan Museum of Zoology and the Illinois Natural History Survey both provided specimens to photograph.

References:

Baker, F.C. 1908. Note on Planorbis binneyi Tryon. The Nautilus 22-23: 41-41.

Burch, J.B. 1980, 1982, 1988. North American Freshwater snails. Walkerana Vol 1(3), 1(4), 2(6). 365 pp.

Dillon, R. T., A. R. Wethington. 2006. No-choice mating experiments among six nominal taxa of the subgenus Physella (Basommatophora: Physidae). Heldia 6: 41 - 50.

Hershler, R. 1994.  A review of the North American freshwater snail genus Pyrgulopsis (Hydrobiidae).  Smithsonian Contributions to Zoology, Smithsonian Institution Press,  115 pp.

Hershler, R.  1996.  Review of the North American aquatic snail genus Probythinella (Rissooidiae: Hydrobiidae).  Invertebrate Biology 115:120-144.

Hershler, R. and F.G. Thompson.  1988  Notes on morphology of Amnicola limosa (Say, 1817) (Gastropoda: Hydrobiidae) with comments on status of the subfamily Amnicolinae.  Malacological Review 21:81-92. 

Hershler, R., J.R. Holsinger, L. Hubricht.   1990.  A revision of the North America freshwater snail genus Fontigens (Prosobranchia:Hydrobiidae).  Smithsonian Contributions to Zoology, Smithsonian Institution Press, 49pp.

Hershler, R., L. Hsiu-Ping, F.G. Thompson.  2002.  Phylogenetic relationships of North American nymphophiline gastropods based on mitochondrial DNA sequences.  Zoologica Scripta 32:357-366.

Pip, E. and J.P.C. Frank. 2008. Molecular phylogenetics of central Canadian Physidae (Pulmonata: Basommatophora). Canadian Journal of Zoology 86: 10–16.

Turgeon, D.D., J.F. Quinn, Jr., A.E. Bogan, E.V. Coan, F.G. Hocherg, W.G. Lyons, et al. 1998. Common and Scientific names of aquatic invertebrates from the United States and Canada: Mollusks, 2nd ed. pp. 526.

Walther, A.C., J.B. Burch, D. O Foighil. 2010. Molecular phylogenetic revision of the freshwater limpet genus Ferrissia (Planorbidae: Ancylinae) in North America yields two species: Ferrissia (Ferrissia) rivularis and Ferrissia (Kincaidilla) fragilis. Malacologia 53(1):24-45.

Wethington, A. R. , C. Lydeard. 2007. A molecular phylogeny of Physidae (Gastropoda: Basommatophora) based on mitochondrial DNA sequences. Journal of Molluscan Studies 73: 241-257.

 

 

Errors, comments, concerns, or suggestions please send to kperez@uwlax.edu